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Significantly, as shown in Figure 6 in their Extended Data (9), Gordon et al

In detail, at the COP1 WD40a -DET1 interface, the guanidinium group of DET1 R488 coordinates with the side chains of COP1 E492/T494

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This assay employs a proprietary non-fluorescent and pH-dependent glutathione sensor that becomes strongly fluorescent upon reacting with thiol